Application Guidelines
Please use the on-line application to apply to any one of the ten doctoral programs listed. Note that you have the option of indicating one additional program. This program may consider your application if you do not receive an offer of admission from your primary selection.

We ask that we receive your application by December 8, 2007. Although we will evaluate any application received after this date, it is to your advantage to submit your application by the deadline, even if it is not complete at that time.

Supporting Documents:
Transcripts, letters of evaluation, and test scores that do not accompany the application must be sent directly to the Office of Graduate Affairs, 701 West 168th St., Room 205, New York, NY 10032. Please indicate on the outside of the envelope the name of the primary doctoral program to which you are applying.

Test Scores:
To direct GRE and/or TOEFL scores to us, specify Institution Code 2619 on your Education Testing Service (ETS) forms. No department code is necessary. If you report TOEFL scores on the cover page of the application, please write "P" (paper) or "C" (computerized).

Other Programs:
Biomedical Informatics is the only program that, in addition to a Ph.D. program, also offers a terminal Master's program. This program is not part of the Coordinated Doctoral Program. However, the on-line application should also be used to apply to this Master's Program.

To apply for admission to the M.D./Ph.D. program, do not use the on-line application form. Please obtain an application for this program from the Admissions Office of the Faculty of Medicine, College of Physicians & Surgeons, 630 W. 168th St., New York, NY 10032 (phone, 212-305-3595).

Applications and information for all other doctoral programs of the Graduate School of Arts and Sciences that are not at located at the College of Physicians & Surgeons should be obtained from 107 Low Library, Columbia University, New York, NY 10027 (phone, 212-854-4737; fax, 212-854-2863;

Additional Information:
The names and addresses of directors of the individual doctoral programs are below. For additional information you may contact them directly. For assistance from the Office of Graduate Affairs, contact us by phone (212-305-8058), fax (212-305-1031), or e-mail (

Graduate Program Directors
Biochemistry and Molecular Biophysics
Oliver Hobert (212) 305-3885
Biomedical Informatics
George Hripcsak (212) 305-5712
Cell Biology and Pathobiology
Ronald Liem
(212) 305-4078
Genetics and Development
Virginia Papaioannou (212) 305-4753
Integrated Program
Ronald Liem (212) 305-4078
Lorraine Symington (212) 305-4793
Howard Shuman (212) 305-6913
Neurobiology and Behavior
Darcy Kelley (212) 854-5108
Carol Mason (212) 305-2105
Ken Miller (212) 543-5238
Debra Wolgemuth (212) 305-7900
Daniel Goldberg (212) 305-1673
Physiology and Cellular Biophysics
Amy MacDermott (212) 305-3889
John Loike (212) 305-1510